Scotland and France have each called two players into their squads for the final weekend of the Six Nations, after suffering injury setbacks in their respective fourth matches.
Gregor Townsend's Scots were hampered by a host of injuries in an 18-11 defeat to Wales on Saturday, with their starting back three of Blair Kinghorn, Tommy Seymour and Darcy Graham all forced off in a bruising contest.
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Kinghorn (ankle) and Seymour (rib) have now been ruled out of their side's clash with England at Twickenham, along with fellow back Stuart Hogg, who remains unavailable due to the shoulder injury he sustained against Ireland last month.
As a result, Ruaridh Jackson and the uncapped Kyle Steyn, both of Glasgow Warriors, have been added to the Scotland squad, while center James Johnstone and forwards Luke Crosbie, Adam Ashe and Tim Swinson have all been released back to their clubs.
France saw Jefferson Poirot and Wenceslas Lauret hurt in its loss to Ireland.
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La Rochelle prop Dany Priso and Montpellier back-row Kelian Galletier have taken their places in the squad for a final-round meeting with Italy.
#XVdeFrance : Après #IRLFRA, @Jeff_Poirot (@UBBrugby) et @lauretwenceslas (@racing92) sont blessés et forfaits pour #ITAFRA. Ils sont remplacés par @Danypriso (@staderochelais) et @Kel_Galletier (@MHR_officiel) qui rejoindront le groupe à Dublin dans la soirée. #NeFaisonsXV pic.twitter.com/5635kGvuQl
— Fédération Française de Rugby (@FFRugby) March 11, 2019